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The hydroxyl radical (OH) is the most damaging ROS due to its extraordinarily high oxidation potential (2.80 V), near diffusioncontrolled reaction kinetics (10 9 10 10 M 1 s 1 ), and indiscriminate reactivity with all biomoleculesinitiating DNA strand breaks via preferential abstraction of the 5 H atom in deoxyribose to form mutagenic lesions like 8hydroxyguanine, triggering lipid peroxidation cascades that yield cytotoxic aldehydes (e.g., malondialdehyde), and oxidizing amino acid residues (e.g., Tyr, Cys) to inactivate enzymes and promote protein aggregation [27, 28]